Echidna



Echidna is a giant serpent demoness that has taken over the Mitis Forest, where she flies around and scatters her Chimera seed-eggs to propagate further demons, with which she plans to create a world where demons would prosper. Both Nero and Dante run into Echidna and fight her during the course of Devil May Cry 4.

Design
In her dragon form, Echidna resembles a giant serpent with feathery frills on her neck, and several eyes on her head. However, the head can open up into four petals to reveal her true body, a woman's torso growing out of the tail, similar to a mermaid, with the serpent's "tongue" splitting large red plumes that grow from her head like hair.

Devil May Cry 4
Nero encounters Echidna while he travels through the forest searching for a way to the Order's headquarters. He is soon ambushed by Echidna while crossing the Ruined Valley, but manages to escape as she destroys the walkway behind him. He finds Echidna again at her den by the Hell Gate, and they fight. Nero manages to defeat Echidna, and grabs her tail with his Devil Bringer to prevent her from escaping into the Hell Gate. However, Echidna wrenches free, leaving Nero with only the Sephirothic Fruit.

On his way back to Fortuna, Dante encounters Echidna laying her egg-seeds throughout the forest. Dante responds by stylishly kicking several of them into the forest. They ricochet off the trees in a complex manner that leads to them all striking Echidna in the face. Furious, she tries to devour Dante, but he forces his way out of her mouth and fights her. In the end, she is killed by Dante, who then uses the Hell Gate's Devil Arm, Gilgamesh to destroy the monument.

Background
Echidna is named after the Greek mythological monster of the same name, whose name was Greek for "she-viper". She was the mother of many monsters, including Cerberus and the Chimera.