Key Items

Key Items are collectible objects that are usually used as keys to special doors or solutions to puzzles, though in Devil May Cry 4 they provide Nero and Dante with new abilities needed to progress through the missions. They are an important part of progressing through a mission, and must be obtained even if they have been obtained on previous playthroughs. All key items, excluding Proud Souls, are finite and can only be obtained by finding it within the appropriate mission.

Pride of Lion

 * (Shadow)

Wing Talisman
The Wing Talisman (羽のタリスマン) is found within a red pillar in the Spiral Well. It is a crystal talisman containing an angel's wing that can be used to activate certain equipment which will propel Nero or Dante upwards if stepped upon. These pads are usually red, though there are some blue ones that effect a much greater response. There is also an elevator within Experiment Disposal which requires the Wing Talisman to be placed inside of it to move, and Nero does so while searching the H.Q. for Kyrie. Dante later retrieves the talisman.

Key of Cronus
The Key of Cronus (クロノスの鍵) is found within a violet pillar in the Key Chamber. It is a magical key intended for use with the Chrono Slicers, time suspension devices being developed by the Order's technology department. Most of them are incomplete and will only remain active for a short time. However, there is one Chrono Slicer within the Security Corridor which functions indefinitely if the key is left inside, and Nero does so while searching the H.Q. for Kyrie. Dante later retrieves the key, and uses it to wreck some of the machines in the Security Corridor.

Background
In Greek mythology, Cronus is the father of Zeus, and the god of "human time" (harvests, calendars, etc.). He is often conflated with Chronos, the divince personification of time itself.