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Mission 02: Highway Star" is the second episode of Devil May Cry: The Animated Series. It aired on June 21, 2007.

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Three racers race their motorcycles at incredibly high speeds, enjoying themselves as they do. While they do, two unknown men in black suits watch them from afar. As the race goes on, one of the racers tells his brother, Michel, that he's driving way too fast. However, his brother rebukes him and tells him he may slow down if he's too scared. As he speeds up, a motorcycle quickly passes him. Feeling challenged, he races the mysterious motorcycle but ends up dead in a crash.

Meanwhile, a mysterious woman stands idly in the rain. A man approaches the lady, offering to find her a warm place and buy her dinner after, assuming she's waiting for her boyfriend. The woman agrees to go with the stranger. The woman leads the man into an abandoned building. The man questions the woman on what's going on, stating there's nothing here of interest. However, the woman finds a closet full of human skulls. The man questions her on what's going on. The woman reveals that he should know and accuses him of being responsible for the killings as she recounts how several women were mysteriously picked up by a man who treats to them dinner and spends a night with them in a hotel. Eventually, the women went missing. The man, seeing the ploy isn't working, then transforms into his true demon form and attacks the woman, who's revealed to be Lady. After Lady kills the demon, she is approached by two unknown men in suits that claim to be part of the Highway Department. They reveal they were looking for Lady, offering her a job.

In the Devil May Cry HQ, Patty complains about the "style" and uncleanliness of the place, while Dante reads peacefully and ignores her complaints. He is then promptly called to collect his pizza from Andy. Patty questions Dante if clients will ever appear. Dante assures that clients wouldn't want to visit his place based on the reaction he got from Andy. It is then Lady arrives at the HQ, commenting on the "style" of the place having been improved since Patty has added her own decorations. Patty takes an immediate liking to Lady as she reveals she knows who Patty is. However, Dante questions the reason for her appearance and inquires if she's attempting to give him a job. He expresses he doesn't want to accept any jobs from her on account that she gives him terrible jobs. She then claims she's here to collect the debt he owes her, but instead, then offers a bet: If she's able to shoot the eight ball into the hole in the pool table, he must take the job unconditionally and if she loses, she must forget the debt and pay him $20,000 in cash. Dante loses the bet and accepts the job.

Arriving at a biker lot, Dante then changes his mind on account that when he deals with humans, he doesn't allow himself to injure humans, making them more irritating to deal with then actual demons. Lady then decides to talk to the bikers herself. She confronts one of the members and requests to see their leader. Vincent then appears, introducing himself as the leader. Lady warns that they should "hit the road" and quit racing, but her warning falls short. Vincent then tells them to stay out of his way, wanting to lure out the racer who killed his brother, Michel. Dante then calls him an idiot for wanting to do so, which angers Vincent. Lady then challenges Vincent to race Dante, telling Vincent it's the best way to lure out the racer known as "Red Eye".

Both Vincent and Dante get ready to race. As they ready themselves, Dante gives Rebellion to Lady since he says he won't be needing it to race. When the race starts, Dante takes the lead, causing Vincent to use nitro to speed past him, which lures out Red Eye. Vincent starts to hallucinate thanks to Red Eye, thinking he sees his brother. After unsuccessfully trying to stop Vincent from chasing after his hallucination, Dante collapses the bridge under them when he hears a mysterious sound. Lady, who has been following Dante, shoots Vincent's tire, causing him to crash, and tosses Dante his sword. Dante surmises that she really called him because she wasn't able to take Red Eye down on her own. Dante manages to stop Red Eye, who then reveals his true form. Lady arrives and kills the demon with the Kalina Ann. With the demon killed, Vincent arrives to see the biomass of Red Eye that once resembled his brother collapse, in which he responds by calling out to his brother emotionally and cradling pieces of the biomass.

Later, back at the DMC HQ, Patty one again complains about the appearance of the place and the fact there's no clients. Dante expresses that he doesn't want any clients at the moment anyways and that he should be getting paid soon once Lady sorts out their pay with the Highway Department. At that moment, Lady calls, revealing that while Dante was paid, he won't be getting the money as his pay was deducted due to the collapse of the bridge and that the remaining money will be used to help repay part of his debt to her. Vincent then arrives at the HQ to thank Dante and challenges him to a race. Dante bets that if he can sink the eight ball into one of the holes of the pool table, Vincent has to buy him dinner, but if he doesn't, Dante will race him.


  • This episode marks the first appearance of Lady in the anime series. 
  • In this episode it's revealed Lady is regularly contracted by humans and various agencies (namely the police) to perform various amount of jobs pertaining to appearances of demons.
  • This episode reveals that Dante is in a heavy amount of debt to Lady. Though how much he owes is unknown, he did say that If he won the bet, he'd be payed $20,000 in cash.
  • Highway Star could be a reference to the song of the same name made by Deep Purple.